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Costolo has not flinched in pruning and reshaping hismanagement team, while Twitter, the company, has been ruthlessin cutting off the smaller companies that were once a part ofits orbit. A one-time comic actor who cut his teeth in businessat Andersen Consulting before starting several companies,Costolo may never be as closely associated with Twitter as MarkZuckerberg is with Facebook, yet he is arguably just asimportant.